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Age-calibration of hominin fossils and artifacts in the East African Rift is principally achieved through dating of associated volcanic-sedimentary strata. The dominant dating techniques for sites ≳100 ka are the 40Ar/39Ar radiometric dating method, magnetostratigraphy, and tephrostratigraphy. The 40Ar/39Ar technique relies on the occurrence of volcanic deposits in the target strata, which are often present as a consequence of the interplay of rift formation and volcanic activity. The frequency of datable material may be limited; however, by also applying the relative dating methods of magnetostratigraphy and tephrostratigraphy to these same strata, a chronostrati-graphic framework can be built and applied to fossiliferous strata. This chapter provides examples of the application of these techniques at Olduvai Gorge, Tanzania, and two areas of the East African Rift in Ethiopia.more » « less
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East Africa is a global biodiversity hotspot and exhibits distinct longitudinal diversity gradients from west to east in freshwater fishes and forest mammals. The assembly of this exceptional biodiversity and the drivers behind diversity gradients remain poorly understood, with diversification often studied at local scales and less attention paid to biotic exchange between Afrotropical regions. Here, we reconstruct a river system that existed for several millennia along the now semiarid Kenya Rift Valley during the humid early Holocene and show how this river system influenced postglacial dispersal of fishes and mammals due to its dual role as a dispersal corridor and barrier. Using geomorphological, geochronological, isotopic, and fossil analyses and a synthesis of radiocarbon dates, we find that the overflow of Kenyan rift lakes between 12 and 8 ka before present formed a bidirectional river system consisting of a “Northern River” connected to the Nile Basin and a “Southern River,” a closed basin. The drainage divide between these rivers represented the only viable terrestrial dispersal corridor across the rift. The degree and duration of past hydrological connectivity between adjacent river basins determined spatial diversity gradients for East African fishes. Our reconstruction explains the isolated distribution of Nilotic fish species in modern Kenyan rift lakes, Guineo-Congolian mammal species in forests east of the Kenya Rift, and recent incipient vertebrate speciation and local endemism in this region. Climate-driven rearrangements of drainage networks unrelated to tectonic activity contributed significantly to the assembly of species diversity and modern faunas in the East African biodiversity hotspot.more » « less
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